International Database for Barley Genes and Barley Genetic Stocks

BGS 229, Curly dwarf 2, cud2

BGN  47:103
Stock number: BGS 229
Locus name: Curly dwarf 2
Locus symbol: cud2

Previous nomenclature and gene symbolization:



Monofactorial recessive (3).
Located in chromosome 1HL (3); cud2.b is about 14.6 cM proximal from the Blp1 (Black lemma and pericarp 1) locus (3, 4); no SNP markers different than those of Bowman were retained in the Bowman backcross-derived line BW199 for cud2.b (1).


Lemmas and awns are curly or strongly twisted, and stem internodes are curved. Culms are short (1/2 normal), spikes are semi-compact, awns are short (2/3 normal length), and kernels are globe-shaped. Most leaves are short and slightly twisted (3). In the Bowman backcross-derived line for cud2.b, BW199, plants were short (3/5 of Bowman), the twisted leaf blades were slightly shorter, peduncles were coiled and about 1/2 normal length, rachis internodes were slightly shorter, and awns were twisted and 2/3 normal length. Compared to Bowman, kernels were short (80% as long) and slightly wider and were about 3/4 normal weight (4.2 vs. 5.5 mg), and test weights were about 25% lower. Grain yields of BW199 were 1/4 to 1/2 of those for Bowman (2).

Origin of mutant:

An ethyl methanesulfonate induced mutant in Akashinriki (OUJ659, PI 467400) (3).

Mutational events:

cud2.b (OUM112, GSHO 1712) in Akashinriki (OUJ659, PI 467400) (3).

Mutant used for description and seed stocks:

cud2.b (OUM112, GSHO 1712) in Akashinriki; cud2.b in Bowman (PI 483237)*2 (GSHO 2062); cud2.b in Bowman*7 (BW199, NGB 22031).


1. Druka, A., J. Franckowiak, U. Lundqvist, N. Bonar, J. Alexander, K. Houston, S. Radovic, F. Shahinnia, V. Vendramin, M. Morgante, N. Stein, and R. Waugh. 2011. Genetic dissection of barley morphology and development. Plant Physiol. 155:617-627.
2. Franckowiak, J.D. (Unpublished).
3. Hayashi, J., T. Konishi, I. Moriya, and R. Takahashi. 1984. Inheritance and linkage studies in barley VI. Ten mutant genes located on chromosomes 1 to 7, except 3. Ber. Ohara Inst. landw. Biol., Okayama Univ. 18:227-250.
4. Jensen, J. 1986. Coordinator's report: Chromosome 5. Barley Genet. Newsl. 16:32-34.


T. Konishi and J.D. Franckowiak. 1997. Barley Genet. Newsl. 26:227.


J.D. Franckowiak. 2014. Barley Genet. Newsl. 44:111.
J.D. Franckowiak. 2017. Barley Genet. Newsl. 47:103.