BGS 355, Glossy sheath 5, gsh5

Glossy sheath 5 = gs5 (15).
Eceriferum-s = cer-s (5, 13).
Glossy sheath 8 = gs8 (17).

Monofactorial recessive (5).
Located in chromosome 2HL (15, 16, 17, 19); gsh5 is located in the centromeric region about 2.5 cM proximal from the eog1 (elongated outer glume 1) locus (15, 17); gsh5.m is about 32.0 cM proximal from the vrs1 (six-rowed spike 1) locus (15, 17); gsh5.m is associated with SNP markers 1_0851 to 2_1437 (positions 81.43 to 113.28 cM) in 2H bins 06 to 08 of the Bowman backcross-derived line BW408 (2); cer-s.31 is associated with SNP markers 1_0796 to 2_0528 (positions 95.53 to 118.78 cM) in 2H bins 07 to 08 of the Bowman backcross-derived line BW122 (2), in 2H bin 07.

Mutants have a reduced surface wax covering on the spikes and stems, and a bright glossy green color on all leaf sheaths (15) (wax code + - ++) (5). The nodes appear to have an excessive wax covering (4). The Bowman backcross-derived line with the cer-s.31 allele, BW122, headed 2 to 4 days later than BW408 with the gsh5.m allele and Bowman (4). BW122 also had 3 to 6 more kernels per spike. Kernels of BW122 and BW408 were about 10% lighter than those of Bowman. The grain yields of BW408 were often slightly lower than those of Bowman or BW408 (4). The differences in maturity and number of kernels per spike between BW122 and BW408 were attributed to close linkage between alleles at the gsh5 locus and those at the [Eam6 [Early maturity 6, Earliness per se 2S (eps2S), Praematurum-c (mat-c), Hordeum vulgare CENTRORADIALIS (HvCEN)] locus (1, 4).

An X-ray induced mutant in Bonus (NGB 14657, PI 189763) (5); a radiation induced mutant in Jotun (PI 467357) (15); a spontaneous mutant in Okaiku 3 (OUL011) (18).

cer-s.24 (NGB 110908) in Bonus (NGB 14657, PI 189763) (5); cer-s.31 (NGB 110915, GSHO 440) in Bonus (3, 5); cer-s.41 (NGB 110925) in Bonus (5); cer-s.287 (NGB 111174), -s.303 (NGB 111190), -s.372 (NGB 111259), -s.444 (NGB 111332) in Foma (NGB 14659, CIho 11333) (13); cer-s.1047 (NGB 111935) in Carlsberg II (NGB 5085, CIho 10114) (6); cer-s.137 (NGB 111022) in Bonus, -s.591 (NGB 111479) in Foma, -s.803 (NGB 111691) in Bonus, -s.1132 (NGB 112020) in Kristina (NGB 14661, NGB 1500) (7); cer-s.914 (NGB 111802) in Bonus (8); cer-s.622 (NGB 111510) in Bonus, -s.1054 (NGB 111942) in Carlsberg II, -s.1153 (NGB 117264) in Kristina (9); cer-s.1323 (NGB 112211) in Bonus (10); cer-s.1134 (NGB 112022) in Kristina (11); cer-s.1754 (NGB 112568) in Bonus (12); gsh5.m (GSHO 739) in Jotun (PI 467357) (15); gsh5.n (GSHO 832) in Okaiku 3 (OUL011) (14, 18).

gsh5.m (GSHO 739) in Jotun; cer-s.31 (NGB 110915, GSHO 440) in Bonus; cer-s.31 in Bowman (PI 483237)*5 (GSHO 1888); cer-s.31 in Bowman*8 (BW122, NGB 20528); gsh5.m in Bowman*8 (GSHO 1889, BW408, NGB 20641).

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4. Franckowiak, J.D. (Unpublished).
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W.L. McProud and R.F. Eslick. 1971. Barley Genet. Newsl. 1:171-172.

J.D. Franckowiak and U. Lundqvist. 1997. Barley Genet. Newsl. 26:300-301.
U. Lundqvist and J.D. Franckowiak. 2011. Barley Genet. Newsl. 41:149-150.
U. Lundqvist and J.D. Franckowiak. 2014. Barley Genet. Newsl. 44:126-127.
U. Lundqvist and J.D. Franckowiak. 2017. Barley Genet. Newsl. 47:128-130.